The effect of contrasting crown attributes in clonal Pinus taeda trees on forest growth, stand dynamics, and biomass distribution

Objectives

The primary objective of this study is to examine how the differing crown attributes produced by three clonal varieties and one family of loblolly pine (Pinus taeda) affect forest growth, biomass distribution, and stand dynamics. Here we report the contrast for two Clones: A and B

A secondary objective is to examine how these crown attributes correspond to the thinning response of the different genotypes.

Summary / Lessons Learned / Additional Thoughts

Two clones differed in their crown characteristics. Clone B added more height and had a cleaner (no branches) bole under narrow spacing than clone A. This suggests genetics could be used to select for desirable traits but the effect may depend on tree spacing.

Future reporting will include updates on the thinning response of the two clones. Clone A has a larger crown than Clone B, suggesting it will respond more to thinning.

Long Description of Practice

The previous slash pine plantation (established approximately in 2002) was whole-tree harvested in 2009. Prior to this harvest, the site was treated with glyphosate herbicide (Makaze, 5.2 dm3 ac-1, 2 quart ac-1) with a surfactant (Li700, 0.585 l ha-1, 8 oz ac-1).

The trees were treated with 36.8% permethrin at a rate of 1.9 liters (2 quarts) per 100,000 individuals just prior to planting. The study site was hand-planted with containerized seedlings in February 2010. Genetic entries consisted of three loblolly pine clones (Clone A, Clone B, and Clone C) derived from somatic embryogenesis, and a loblolly pine full-sib seedling family (Family B) (see notes on data collection process below for plot design specifics).

For the three years after planting, weed competition was treated as needed with glyphosate (5%), sulfometuron methyl (Sulfomet XP, 0.11 l ha-1,1.5 oz ac-1), clopyralid Sp (Transline, 1.17 l ha-1, 16 oz ac-1), and mowing. This kept the understory relatively free of vegetation while the overstory developed. In April 2010, trees were treated with fipronil (PTM, 1 ml chemical + 9 ml water per tree injected into root zone) to protect against infestation by Nantucket pine tip moth (Rhyacionia frustrana (Comstock)). In May 2010, any dead trees were replaced with the same genotype from extra trees stored on site in container racks. In September 2011, the entire experiment was broadcast fertilized with 535 kg ha-1 of a 10-10-10 + micronutrient blend with 0.05% Mn, 0.07% Zn, 0.17% Fe, 0.02% Cu, and 0.02% B.

Tree Diameters Table

Date (month / year)Species

Tree Heights Table

Date (month / year)Species

Volume Table

Date (month / year)Species

Other Table

Date (month / year)SpeciesCloneSpacing (ft) Narrow=6 x 12 Wide = 12 x 12DBH (in)Height (ft)Ht. to Base of Live crown
3/2021 (age 10)Loblolly pineANarrow7.347.924.0
3/2021 (age 10)Loblolly pineAWide8.651.224.0
3/2021 (age 10)Loblolly pineBNarrow7.259.426.2
3/2021 (age 10)Loblolly pineBWide8.246.920.3

Study Location / Installation Information

Lattitude

29.721697

Longitude

-82.417938

Accessibility

Landowner HAS consented to property visits WITH PERMISSION

Who installed study

First name: Tim

Last name: Martin

 

Year of installation

2010

Does study have end date

NO

Planned or actual end date

Information not provided.

Contact for access

Is study monumented

NO

Stand Information Pretreatment

Land Use History

The study site was previously a slash pine (Pinus elliottii) plantation that had been growing for 7 years before being whole-tree harvested in 2009. That was itself preceded by at least 10 years of horticultural crop experiments.

Stand Area

Information not provided.

Stand Age Before Treatment

7 Years Old

Pretreatment Overstory Community Composition

Slash pine (Pinus elliottii)

Pretreatment Forest Metrics

Total Basal Area

Information not provided.

Species Composition

Information not provided.

Soil Series

Pomona (Spodosol), Millhopper (Ultisol), and Pelham (Ultisol) series. While the Pomona and Pelham series are classified as poorly drained, and Millhopper well-drained, the site has been ditched and there has been no indication (e.g., standing water, ground vegetation) of differences in soil moisture across the study area.

Silvicultural Application(s) Used in Study

What did the study investigate?

  • Other

Other Investigations

Genetics (Clonal and family variation)

If silviculture included a harvest, what type(s)?

Information not provided.

Silvicultural Action(s) Timings and Types / Intensities

Information not provided.

Are there any local forest health threats to your target species?

Yes

Notes on silvicultural process(es) or data collection

The plots were designed as follows: split-split plot, randomized complete block with three replicates. The first split plot was between a thinning treatment (to-be-thinned or no-thin), and then each of those subplots was split between two planting spacing treatments (1.83 m x 3.66 m = 1,493 trees ha-1, and 3.66 m x 3.66 m = 747 trees ha-1). Within each split-split plot were five plots containing the genetic entries: three monoclonal plots, each containing one of three clones, a polyclonal plot containing the three clones randomly distributed, and a full-sib family plot. Plot sizes were designed to vary with spacing and thinning treatment so that all plots would have similar numbers of trees after thinning. Plot sizes were no-thin narrow spacing, 71 trees, 0.05 ha; to-be-thinned narrow spacing and no-thin wide spacing, 144 trees and 72 trees, respectively, 0.10 ha; and to-be-thinned wide spacing, 121 trees, 0.16 ha. In total, approximately 6,000 trees were planted across 60 plots. During the time reported in this manuscript, the “to-be-thinned” treatment remained unthinned.

Genetic entries consisted of three loblolly pine clones (Clone A, Clone B, and Clone C) derived from somatic embryogenesis, and a loblolly pine full-sib seedling family (Family B). Clone B was derived from a progeny of the Family B cross. Clones B and C shared a parent in common. Clone A was unrelated to the other two clones or to the full-sib family. The propagules were grown in containers in the Plum Creek nursery in Jesup, Georgia.

Plots varied in size depending on which split/subsection they were in. Planting densities also varied. See attached paper for specifics.

See attached paper for data collection specifics.

Plans for Future Treatments

'To-be-thinned' plots were thinned. This data will be added to a future report.

Only the significant contrast for Clone A and B are shown in this report.

Plans for Future Measurements

Future measurements in this trial will enable us to examine how different varieties respond to thinning; how previously-observed differences in crown dimensions and growth responses are affected by thinning; whether different varieties begin self-thinning at different relative densities; and other stand dynamics and competition topics.

One prediction associated with narrow-crowned crop ideotypes is that they might have increased carrying capacity or stockability, possibly realized through increased individual tree resource efficiency and an associated potential to carry higher relative density (At age 10 years, the narrow spacing treatments in the current study had just reached relative density consistent with the onset of density-dependent mortality). The study was subsequently thinned at age 11, and so at the time of writing contains a wide range of relative density conditions, from free-to-grow to experiencing density-dependent mortality, conditions favorable for testing the stockability of clones across this gradient.

Costs and Economic Considerations

These were not tracked as it was an experiment, but the number of treatments associated with this installation would make it on the higher end of cost for a plantation. One cost savings is planting in an old-field which reduced the site preparation costs.

Scientific Study Information

Is this study published?

No

Study Design

Split-split plot, randomized complete block

Number of Replicates

3

Size of Plots

3 Acres

Planting Density

Information not provided.